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Spanish toothcarp

Apricaphanius iberus

Fishes › Killifish and livebearers (Cyprinodontiformes) › Aphaniidae (Aphanius killifish)

The Spanish toothcarp (Apricaphanius iberus), also known as the Spanish pupfish or Iberian killifish, is a small species of ray-finned fish in the family Aphaniidae endemic to Spain.

Intro from Wikipedia (CC BY-SA 4.0)

Aquarium hobby
Experience: Intermediate
Spanish toothcarp
Photo: no rights reserved, CC0, via iNaturalist
FamilyAphanius killifish
Adult lengthup to 2.2 in
WaterFreshwater, brackish
HabitatLives in freshwater and brackish water

Care at a glance

Species-specific aquarium care

  • Tank: At least 24 in long and 12 in front to back for one adult (adults commonly reach about 2.2 in / 5.5 cm).
  • Water: Freshwater (tolerates brackish).
  • Temperature: 50–90°F (range recorded for this species).
  • pH: 6.5–7.5.
  • Hardness: 8–10 dGH.
  • Layout: Uses both the bottom and open water: give open swimming space plus cover near the substrate.
  • Diet: A predator: feed meaty foods sized to its mouth (frozen fish, shrimp, worms); it will eat tankmates that fit in its mouth. FishBase classes it as a carnivore.

Calculated from this species' own FishBase record: tank size from its typical adult length and swimming zone, water values from its recorded ranges. The group guide below fills in general care.

Care guide

Species care notes

  • Tank: a base of at least 47 × 12 in (roughly 29 US gallons); space needed is rated “large tank”.
  • Water: a fairly wide range, from slightly acidic to slightly alkaline, 36–86 °F, pH 7–9, 10–30 dGH.
  • Behaviour: a shoaling fish best kept in a group of six or more.
  • Difficulty: Moderate (Seriously Fish rating).

Short excerpts from the species profile:

  • Setup: “Even for long-term maintenance a simple set-up will suffice.”
  • Diet: “Apricaphanius species are basically micropredators feeding on small aquatic crustaceans, worms, insect larvae and other zooplankton although algae and other plant material is also taken at times.”
  • Tankmates: “Its particular water requirements and aggressive spawning behaviour make A.”

From the Seriously Fish profile for this species, which has the full habitat, maintenance, diet, behaviour and breeding write-up 167

Group care guide

Aphanius killifish

Applies to most species in this group. Habitat: These toothcarps occupy vegetated Mediterranean coastal lagoons and inland Anatolian springs, streams and lake margins with locality-dependent salinity (fasciatus, anatoliae). Individual species can differ, so compare with the species numbers above and the linked sources.

The published 35.6–86°F seasonal envelope for fasciatus/anatoliae is not a constant setpoint or a family-wide recommendation. Aphaniops dispar/sirhani, Apricaphanius iberus and the restricted Anatolichthys species need their own locality protocols; keep provenance and avoid mixing conservation lines. Some toothcarps are legally protected (fasciatus, anatoliae). Level is an editorial assessment.

Housing & setup

  • For fasciatus/anatoliae groups use a footprint of at least 120 × 30 cm; fasciatus pairs/trios can use 60 × 30 cm (fasciatus, anatoliae).
  • Fill substantial space with spawning mops or fine-leaved plants to break male sightlines (fasciatus, anatoliae).
  • Provide hard alkaline water: fasciatus pH 7.5–9.0, anatoliae 7.0–8.5; both 10–30 dGH (fasciatus, anatoliae).
  • Set salinity from collection locality, rather than treating every population as freshwater (fasciatus, anatoliae).
  • Use gentle filtration for anatoliae and preserve a seasonal cool winter period (anatoliae).

Sources 1127112811291130

Diet

  • Live or frozen Artemia and Daphnia (fasciatus, anatoliae).
  • Bloodworms (fasciatus, anatoliae).
  • Spirulina-containing dried food and filamentous algae (fasciatus, anatoliae).

Sources 1127112811291130

Social life

  • Maintain species/locality breeding groups, with two or three females per male for anatoliae and refuges from dominant males (anatoliae).

Sources 1127112811291130

Health watch

  • Ammonia toxicity from inadequate biofiltration (fish_health).
  • Parasitic skin and gill infections (fish_health).
  • Nutritional deficiencies from unsuitable diets (fish_health).

Sources 1127112811291130

Understanding the animal

In the wild

In the wild

From this species’ own FishBase record (habitat, bottom type, water and breeding data), not its family. Good care copies these conditions.1

Where it livesLives in freshwater and brackish water, in rivers and streams, marshes and swamps, estuaries. Uses both the bottom and open water. Inhabits lagoons, salt marshes, swamps, estuaries and freshwaters. Euryhaline, occurring in water bodies with salinities ranging from freshwater to 57 ppt.
Water it lives inFreshwater / brackish water, 50–90 °F (10–32 °C), pH 6.5–7.5, hardness 8–10 dH.
ClimateTemperate climate.
Wild dietHunting macrofauna (predator).
Behavior to plan forBreeding: open water/substratum egg scatterers.

What this means for care

  • Keep water within its wild range, 50–90 °F.
  • It tolerates or needs brackish water; check salinity needs before setup.
  1. FishBase species summary: Apricaphanius iberus
Native climate (measured)

Native climate (measured)

Warm-season highsabout 84°F (up to 87°F in the hottest sampled areas)
Cool-season lowsabout 39°F (down to 38°F in the coldest sampled areas)
Average temperature59–63°F across sampled records
Relative humidity72–72% on average (driest month about 67%)
Rainfallabout 504 mm a year, spread fairly evenly
WintersCool to cold winters (lows around 39°F) in much of its native range. Seasonal slowdown may be relevant, but species-specific biology determines whether brumation or hibernation occurs.

Measured at 5 wild occurrence records sampled against WorldClim 2.1 climate data (1970–2000 averages). Air conditions in the range are ecological context, not enclosure settings: microhabitats such as burrows, leaf litter or basking rocks can be cooler, warmer or more humid.

From the sources

From the sources

Quoted from the matching sections of reputable sources for this exact species. Specialist databases are used first; Wikipedia (community-edited, CC BY-SA 4.0) only fills gaps. Other sources are short credited quotes.

OverviewThe Spanish toothcarp (Apricaphanius iberus), also known as the Spanish pupfish or Iberian killifish, is a small species of ray-finned fish in the family Aphaniidae endemic to Spain. — Wikipedia (community-edited)
Where it livesHome ranges Population dynamics Growth parameters Max. ages / sizes Length-weight rel. Length-frequencies Mass conversion Recruitment Abundance Life cycle Reproduction Maturity Maturity/Gills rel. — FishBase
Wild dietThe Spanish toothcarp is an omnivore, and eats insects, crustaceans, worms and algae. — Wikipedia (community-edited)
ReproductionThe lifespan of a Spanish toothcarp is short, reaching sexual maturity at merely 3 months of age. The females spawn repeatedly each season, producing from 100 to 900 eggs. — Wikipedia (community-edited)
Appearance and sizeThe Spanish toothcarp is a small fish, rarely more than 5 cm in length. Females are longer than males of the same age. It has an oblong body and rounded fins. — Wikipedia (community-edited)
BiologyAdults inhabit lagoons, salt marshes, swamps, estuaries and freshwaters. Euryhaline, occurring in water bodies with salinities ranging from freshwater to 57 ppt. They occur in lowland water with little current. — FishBase
History and namingThe genus Apricaphanius contains three species. One of them, A. saourensis, is found in Algeria, and the other two are endemic to Spain. The other Spanish species is A. baeticus; it was not recognized as distinct from A. iberus until 2002. — Wikipedia (community-edited)
ConservationThe risk of extinction of the Spanish toothcarp is one of the greatest of any Iberian vertebrate. Its limited range, coupled with the drastic population decline the species has suffered in the 1970s, 1980s and 1990s, has caused it to be placed on endangered species lists, both in Spain and internationally. Its conservation status in the south of the Iberian Peninsula has notably worsened. — Wikipedia (community-edited)

Data and studies

The numbers

The numbers

Each bar is what one source reports. Where several sources weigh in, the dashed gold band marks their average.

Adult length

Average of sources2.1in
FishBase 1642.2
Seriously Fish species profiles 1672

Maximum recorded length. Measured as total length. Fish in home aquariums often stay smaller than wild records.

Water temperature

Average of sources43–88°F
FishBase 16450–90
Seriously Fish species profiles 16736–86

Range the species tolerates in nature; aim for the middle.

pH

Average of sources6.8–8.2
FishBase 1646.5–7.5
Seriously Fish species profiles 1677–9

Water hardness

Average of sources9–20dGH
FishBase 1648–10
Seriously Fish species profiles 16710–30

Minimum tank footprint

Reported47in
Seriously Fish species profiles 16747

Seriously Fish recommends a tank base of at least 47 × 12 in (about 29 US gallons).

Sources for this sheet

  1. FishBase (Froese & Pauly, eds.), version 24.07 Peer-reviewed researchThe global scientific database on fishes, maintained by a consortium of research institutions; every value is referenced.
  2. Seriously Fish species profiles Experienced keeper or breederLong-running aquarium reference written and peer-checked by experienced fishkeepers and ichthyologists; each profile covers habitat, tank size, water, diet, behaviour and breeding.
  3. Seriously Fish: Aphanius fasciatus Experienced keeper or breederAdded in the 2026 source-checked care-guide review.
  4. Seriously Fish: Aphanius anatoliae Experienced keeper or breederAdded in the 2026 source-checked care-guide review.
  5. Merck Veterinary Manual: Disorders and Diseases of Fish Zoo, university or veterinaryAdded in the 2026 source-checked care-guide review.
  6. Merck Veterinary Manual: Management of Aquarium Fish Zoo, university or veterinaryAdded in the 2026 source-checked care-guide review.
  7. Wikipedia (species article) Community referenceFree encyclopedia; useful overviews with their own citations. Intro text is quoted under CC BY-SA 4.0.
  8. iNaturalist (captive observations) Community referenceCommunity observation platform run by the California Academy of Sciences and National Geographic; observations marked captive show animals living in homes, zoos and aquariums.